Licence & Citation

The Human Protein Atlas is licensed under the Creative Commons Attribution-ShareAlike 3.0 International License for all copyrightable parts of our database, specifically indicated in the downloadable XML format with 'source="HPA"' and all downloadable files.

While we encourage you to use these resources for your research and commercial purposes, we want to ensure that our content is given proper citation in all cases where it is used.

If you are unsure about how to cite this source or would like written approval for a specific use that is otherwise restricted, please contact us at:

We encourage you to send us a copy of your publication or other work citing these resources using the email address above. Such examples help us document and better understand how our content is being used, and thus guide and support our continued efforts to provide useful open public resources.

Citation guidelines for the Human Protein Atlas

General citation of the Human Protein Atlas

If you cite or display any content, or reference our organization, in any format, written or otherwise, including print or web publications, presentations, grant applications, websites, other online applications such as blogs, or other works, you must follow these guidelines.

  • Include a reference to a Primary publication, or one of the other publications listed under Other relevant publications, as stated below.

Specific citation to images, genes, or data

If you use images, or reference a specific gene, or other data downloaded from the site, in addition to citing the Human Protein Atlas, please also cite the specific image, gene, or data used and the URL that links directly to that information in a manner that will allow a third party to navigate to that image or data on the site.

  • Include image credit: Human Protein Atlas
  • Include citation to the specific image, gene, or data used and the URL that links directly to that information: image/gene/data available from
    Where ## indicates the desired version of the atlas.

Primary publications

UhlÚn M et al, 2015. Tissue-based map of the human proteome. Science
PubMed: 25613900 DOI: 10.1126/science.1260419

Thul PJ et al, 2017. A subcellular map of the human proteome. Science.
PubMed: 28495876 DOI: 10.1126/science.aal3321

Uhlen M et al, 2017. A pathology atlas of the human cancer transcriptome. Science.
PubMed: 28818916 DOI: 10.1126/science.aan2507

Other relevant publications

Uhlen M et al, 2010. Towards a knowledge-based Human Protein Atlas. Nat Biotechnol.
PubMed: 21139605 DOI: 10.1038/nbt1210-1248

Berglund L et al, 2008. A genecentric Human Protein Atlas for expression profiles based on antibodies. Mol Cell Proteomics.
PubMed: 18669619 DOI: 10.1074/mcp.R800013-MCP200

UhlÚn M et al, 2005. A human protein atlas for normal and cancer tissues based on antibody proteomics. Mol Cell Proteomics.
PubMed: 16127175 DOI: 10.1074/mcp.M500279-MCP200

PontÚn F et al, 2008. The Human Protein Atlas - a tool for pathology. J Pathol.
PubMed: 18853439 DOI: 10.1002/path.2440

Integrating our data into a website

When integrating our data into a website, for example, using our XML-files, cite the source in a manner that is clear, accurate and easily discoverable and link to the source. Also, be sure that our content is never displayed in the absence of such citation.