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MAPT

GENERAL INFORMATION

Gene name

 MAPT

Gene description

 Microtubule-associated protein tau

Protein class

 Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
Plasma proteins
Predicted intracellular proteins

Predicted localization

 Intracellular

Number of transcripts

 12
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HUMAN PROTEIN ATLAS INFORMATION

RNA category
TCGA (cancer tissue):Group enriched (breast cancer, glioma, prostate cancer, renal cancer)
HPA (cell line):Cell line enhanced (HDLM-2, SCLC-21H, T-47d)
HPA (normal tissue):Tissue enriched (cerebral cortex)

Protein evidence

 Evidence at protein level

Protein expression
normal tissue

 Cytoplasmic expression mainly in CNS and peripheral nerves.

ANTIBODY IHC RELIABILITY

Data reliability
description

 Antibody staining mainly consistent with RNA expression data.

Reliability score

 Supported based on 3 antibodies.
HPA069524 , HPA069570 , CAB000151
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PROGNOSTIC SUMMARY
Gene product is not prognostic.
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RNA EXPRESSION OVERVIEW
TCGA dataset
RNA cancer category: Group enriched (breast cancer, glioma, prostate cancer, renal cancer)

PROTEIN EXPRESSION
Colorectal cancer
Breast cancer
Prostate cancer
Lung cancer
Glioma

STAINING SUMMARY
HPA069524
HPA069570
CAB000151

GENE INFORMATION

Gene name

 MAPT (HGNC Symbol)

Synonyms

 DDPAC, FLJ31424, FTDP-17, MAPTL, MGC138549, MSTD, MTBT1, MTBT2, PPND, PPP1R103, tau

Description

 Microtubule-associated protein tau (HGNC Symbol)

Entrez gene summary

 This gene encodes the microtubule-associated protein tau (MAPT) whose transcript undergoes complex, regulated alternative splicing, giving rise to several mRNA species. MAPT transcripts are differentially expressed in the nervous system, depending on stage of neuronal maturation and neuron type. MAPT gene mutations have been associated with several neurodegenerative disorders such as Alzheimer's disease, Pick's disease, frontotemporal dementia, cortico-basal degeneration and progressive supranuclear palsy. [provided by RefSeq, Jul 2008]

Chromosome

 17

Cytoband

 q21.31

Chromosome location (bp)

 45894382 - 46028334

Protein evidence

 Evidence at protein level (all genes)

Ensembl

 ENSG00000186868 (version 83.38)

Entrez gene

 4137

UniProt

 P10636 (UniProt - Evidence at protein level)

neXtProt

 NX_P10636

Antibodypedia

 MAPT antibodies

PROTEIN BROWSER
«
MAPT-001
MAPT-002
MAPT-003
MAPT-004
MAPT-006
MAPT-007
MAPT-008
MAPT-201
MAPT-202
MAPT-203
MAPT-204
MAPT-205
»

PROTEIN INFORMATION

Splice variant

UniProt

Protein class

Gene ontology

Length & mass

Signal peptide
(predicted)

Transmembrane regions
(predicted)
MAPT-001
ENSP00000334886
ENST00000334239
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 352 aa
36.8 kDa 		No 0
MAPT-002
ENSP00000408975
ENST00000446361
P10636 [Direct mapping]
Microtubule-associated protein tau
A0A024R9Y0 [Target identity:100%; Query identity:100%]
Microtubule-associated protein
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 383 aa
40 kDa 		No 0
MAPT-003
ENSP00000413056
ENST00000420682
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 412 aa
43 kDa 		No 0
MAPT-004
ENSP00000410838
ENST00000415613
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 776 aa
80.9 kDa 		No 0
MAPT-006
ENSP00000389250
ENST00000431008
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 410 aa
42.6 kDa 		No 0
MAPT-007
ENSP00000460965
ENST00000574436
P10636 [Direct mapping]
Microtubule-associated protein tau
A0A024RA17 [Target identity:100%; Query identity:100%]
Microtubule-associated protein
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 441 aa
45.8 kDa 		No 0
MAPT-008
ENSP00000458742
ENST00000571987
P10636 [Direct mapping]
Microtubule-associated protein tau
A0A024R9Y1 [Target identity:100%; Query identity:100%]
Microtubule-associated protein
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 758 aa
78.9 kDa 		No 0
MAPT-201
ENSP00000262410
ENST00000262410
P10636 [Direct mapping]
Microtubule-associated protein tau
A0A024R9Y1 [Target identity:100%; Query identity:100%]
Microtubule-associated protein
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 758 aa
78.9 kDa 		No 0
MAPT-202
ENSP00000340438
ENST00000340799
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 412 aa
43 kDa 		No 0
MAPT-203
ENSP00000340820
ENST00000344290
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0001764 [neuron migration]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005634 [nucleus]
GO:0005737 [cytoplasm]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0005930 [axoneme]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0007409 [axonogenesis]
GO:0007420 [brain development]
GO:0007565 [female pregnancy]
GO:0007584 [response to nutrient]
GO:0007628 [adult walking behavior]
GO:0008017 [microtubule binding]
GO:0008088 [axon cargo transport]
GO:0008631 [intrinsic apoptotic signaling pathway in response to oxidative stress]
GO:0010033 [response to organic substance]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0014069 [postsynaptic density]
GO:0015630 [microtubule cytoskeleton]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0019901 [protein kinase binding]
GO:0019904 [protein domain specific binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0032387 [negative regulation of intracellular transport]
GO:0032403 [protein complex binding]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0043005 [neuron projection]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0047497 [mitochondrion transport along microtubule]
GO:0048675 [axon extension]
GO:0048699 [generation of neurons]
GO:0051721 [protein phosphatase 2A binding]
GO:0060632 [regulation of microtubule-based movement]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 776 aa
80.9 kDa 		No 0
MAPT-204
ENSP00000303214
ENST00000351559
P10636 [Direct mapping]
Microtubule-associated protein tau
A0A024RA17 [Target identity:100%; Query identity:100%]
Microtubule-associated protein
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 441 aa
45.8 kDa 		No 0
MAPT-205
ENSP00000443028
ENST00000535772
P10636 [Direct mapping]
Microtubule-associated protein tau
Show all
Predicted intracellular proteins
Plasma proteins
Candidate cardiovascular disease genes
Disease related genes
FDA approved drug targets
   Small molecule drugs
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000226 [microtubule cytoskeleton organization]
GO:0005200 [structural constituent of cytoskeleton]
GO:0005515 [protein binding]
GO:0005829 [cytosol]
GO:0005874 [microtubule]
GO:0005875 [microtubule associated complex]
GO:0005886 [plasma membrane]
GO:0006915 [apoptotic process]
GO:0006921 [cellular component disassembly involved in execution phase of apoptosis]
GO:0008017 [microtubule binding]
GO:0010506 [regulation of autophagy]
GO:0012501 [programmed cell death]
GO:0015631 [tubulin binding]
GO:0017124 [SH3 domain binding]
GO:0019899 [enzyme binding]
GO:0030424 [axon]
GO:0030425 [dendrite]
GO:0030426 [growth cone]
GO:0031113 [regulation of microtubule polymerization]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0034185 [apolipoprotein binding]
GO:0034399 [nuclear periphery]
GO:0036464 [cytoplasmic ribonucleoprotein granule]
GO:0044297 [cell body]
GO:0045298 [tubulin complex]
GO:0045773 [positive regulation of axon extension]
GO:0048699 [generation of neurons]
GO:0071813 [lipoprotein particle binding]
GO:0097418 [neurofibrillary tangle]
Show all
 410 aa
42.6 kDa 		No 0

Contact

The Project

The Human Protein Atlas

The Human Protein Atlas project is funded
by the Knut & Alice Wallenberg foundation.